Regarding the green roots on your
Paph spicerianum, I really don't know the answer to this one. I'd have to spot some in-situ pics, or actually find some in the wild to know why they would have green roots.
All I know is that not all Paphs are lithophytic or "terrestrial". Some Paphs are actually epiphytic.
Paph villosum is an example of an epiphytic Paph. The photo evidence is in the book "Wild Orchids of Myanmar, Volume 1: Last Paradise of Wild Orchids" by Dr. Yoshitaka Tanaka, Nyan Htun, and Tin Tin Yee.
Although the text out there I see for
Paph spicerianum says that they either grow as a lithophyte or a terrestrial orchid, it may not be the entire truth.
However, I have seen, (and do own), lithophytes and a few terrestrial orchids that have green roots as well.
Everything I've said so far unfortunately, all comes back to, I don't know why the roots are green.
I can point you to something else interesting...
I found an article that talked about green roots in terrestrial plants that were in the "Sunflower Family", (
Asteraceae) and the "Nightshade Family", (
Solanaceae):
http://www.plantphysiol.org/content/101/2/363.full.pdf
Why is this relevant to orchids?
This article may not be addressing orchids directly, but it does address green roots. Green roots are clearly an evolutionary advantage to some of the largest families of flowering plants in existence. And this may have had something to do with some significant environmental event in their evolutionary past that gave plants with this trait an edge in survival. What that historical environmental event is, I have no clue whatsoever.
In terms of the advantage, it should be evident that the advantage is 2 sources of food production.
1. The leaves.
2. The roots.